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Personal profile


  • 2003-present Professor of Developmental Neurobiology, University of Edinburgh
  • 1996-2003 Reader, University of Edinburgh
  • 1988-1996 Lecturer and Senior Lecturer, University of Edinburgh
  • 1987-1988 MRC Travelling Fellow, University of California at Berkeley, USA
  • 1985-1986 Beit Memorial Research Fellow, Physiology, Oxford University
  • 1982-1985 MRC Training Fellow, Physiology, Oxford University

My research in a nutshell

My team takes a multidisciplinary approach, using a range of neuroanatomical, electrophysiological, cell biological, molecular biological, transplantation, genetic and transgenic methods to analyse mechanisms regulating events that occur during the pre- and post-natal development of cortex in various species. Our results demonstrate that the transcription factor, Pax6, controls cell proliferation, migration and differentiation in the embryonic neocortex, and that it regulates the growth of axons from the embryonic thalamus to the cerebral cortex. Using genetics, transgenics, chimaeras and transplantation methods we have discovered some of the mechanisms regulating these processes.



Undergraduate: Year 3: Mechanisms of Brain Development 3; Year 4: Honours Neuroscience, Honours Physiology, Honours elective in Development and Disease

Postgraduate: MSc by Research Integrative Neuroscience;  MSc by Research Biomedical Sciences (Zhejiang); MSc Biomedical Sciences (Cell Communication and Genes and Disease); Integrated PhD Translational Neuroscience

Research Interests

My research concerns the mechanisms that control early development of the mammalian forebrain.


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