A new genus of metriorhynchid crocodylomorph from the Lower Cretaceous of Germany

Here we describe a new genus and species of metriorhynchid crocodylomorph, Enalioetes schroederi gen. et sp. nov., from the lower Valanginian Stadthagen Formation (Lower Cretaceous) of north-western Germany. Enalioetes schroederi is the most complete and well-preserved Cretaceous metriorhynchid skull known to date, preserving most of the cranium and mandible, the atlas-axis complex and the first postaxial cervical vertebra. The specimen was previously attributed to the coeval enigmatic metriorhynchid Enaliosuchus (a nomen dubium) and, more recently, to Cricosaurus. Although the specific epithet schroederi has been used frequently in the literature, it has never been formally established. Herein, we demonstrate that the new taxon is distinct from all known metriorhynchids by a unique combination of characters including several autapomorphies such as: the lack of bulbous dorsolateral expansion in the posterior nasal cavity; mediolateral distance between the orbital canals being approximately 1.5 times the diameter of the orbital canals; lacrimal with dorsoventrally deep anterior process ventral to the preorbital fossa being equal to or greater than the depth of the jugal anterior process and the anterodorsal process of the lacrimal; and ascending processes at atlas intercentrum extending far dorsally to level of neural canal. Enalioetes schroederi contributes to the sparse global record of Cretaceous metriorhynchids and represents one of the stratigraphically youngest occurrences of the group. It can thus help to enhance our understanding of the metriorhynchid diversity during the Cretaceous Period.